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Family: Ericaceae
whiteveined wintergreen, more...white-vein wintergreen, whitevein shinleaf
[Pyrola aphylla Sm., morePyrola aphylla var. leptosepala Nutt., Pyrola aphylla var. paucifolia T.J. Howell, Pyrola blanda Andres, Pyrola conardiana Andres, Pyrola dentata Sm., Pyrola dentata var. apophylla Copeland, Pyrola dentata var. integra A. Gray, Pyrola pallida Greene, Pyrola paradoxa Andres, Pyrola picta ssp. dentata (Sm.) Piper, Pyrola picta ssp. integra (A. Gray) Piper, Pyrola picta ssp. pallida Andres, Pyrola picta var. dentata (Sm.) Dorn, Pyrola septentrionalis Andres, Pyrola sparsifolia Suksdorf] |
Plants rhizomatous, (0.5-)1.5-4.6 dm. Leaves sometimes highly reduced or absent; petiole 4-25 mm, channeled adaxially, glabrous; blade usually maculate, sometimes not maculate, dull and light green to purplish abaxially, dull or shiny and green to dark green, usually with white tissue bordering larger veins adaxially, ovate or ovate-elliptic to oblanceolate or spatulate, 8-80 × 4-49 mm, coriaceous, base obtuse to attenuate or decurrent, margins entire or denticulate to coarsely denticulate, apex obtuse to acute. Inflorescences 1-2(-3), 4-24-flowered; peduncular bracts 1-5, subulate to narrowly lanceolate, 3-9 × 1-2 mm, membranous, margins entire or erose-denticulate; inflorescence bracts subulate to ovate-lanceolate, usually shorter than or, rarely, longer than subtended pedicels, 3.5-6 × 2-3 mm, herbaceous. Pedicels 3-18 mm. Flowers: calyx lobes appressed or spreading in fruit, green with margins hyaline to white or pinkish, widely deltate to widely ovate, 1.1-3.9 × 1.1-2.4 mm, margins entire or erose-denticulate, apices acute; petals greenish white, white, pink, or reddish, obovate, 4.2-11.2 × 3.1-7.1 mm, margins entire or obscurely denticulate; stamens 5-9 mm; filament base 0.5-0.8 mm wide; anthers 1.8-5.5 mm, apiculations absent or 0.1-0.2 mm, thecae creamy white to pink, tubules orange, 0.4-0.8 mm, gradually narrowed from thecae, lateral walls usually touching for most of their lengths, pores 0.1-0.3 × 0.1-0.2 mm; ovary smooth; style exserted, 4-9 mm; stigma (0.8-) 1-1.6 mm wide, lobes erect. Capsules depressed-globose, 3-6 × 6-9 mm. 2n = 46. Flowering Jun-Aug. Dry, coniferous forests; 400-3000 m; Alta., B.C.; Ariz., Calif., Colo., Idaho, Mont., Nev., N.Mex., Oreg., S.Dak., Utah, Wash., Wyo. E. Haber (1987) concluded that Pyrola picta, P. aphylla, and P. dentata are morphotypes of a single, highly variable species, a finding consistent with seed morphology data compiled by Hiroshi Takahashi (1993). Leafless scapes frequently are found attached to rhizomes bearing leafy shoots (W. H. Camp 1940; Haber 1987). Putative hybrids between P. picta and P. chlorantha have been reported at three locations in the western United States (Haber 1993). Cladistic analyses of molecular and morphologic data suggest that P. picta is sister to P. chlorantha (J. V. Freudenstein 1999b), which also occasionally is leafless.
Plant: Perennial rhizomatous, scapose herb Leaves: blades ovate, elliptic, to oblanceolate, (0.8-)3.5-6.0 (-8.0) cm long, leathery, dark green above, commonly purple below; veins generally white-bordered; margins minutely crenately toothed or denticulate INFLORESCENCE: glabrous symmetric bracteate racemes Flowers: with sepals ovate, these (1.0-)1.5-3.0(-3.9) mm long, the tips acute; petals greenish, cream-white or pink, (5.0-)6.0-7.5(-8.9) mm long; anthers (2.5-)3.0-4.0(-4.5) mm long, the tubes prominent, the pores small Fruit: FRUITS 5-locular drooping capsules with a depressed center and fibers connecting opened valves Misc: Dry coniferous forest; 2400-2900 m (8000-9500 ft); Jul-Aug Notes: scapes (3-)15-25(-36) cm tall; bracts all ovate-lanceolate to subulate, shorter than pedicels REFERENCES: Haber, Erich. 1992. Pyrolaceae. Ariz.-Nev. Acad. Sci. 26(1)2. |