Perennials. Stems pilosulous to villous and/or stipitate-glandular. Leaves: petioles 3-8(-12+) cm, sometimes raggedly winged; blades 4-20(-40+) × 2-12(-25+) cm, larger usually deeply pinnately lobed, lobes 5-7. Cypselae 3-4 mm, 3-angled or -ribbed. 2n = 30. Flowering May-Aug(-Oct). Damp, shaded sites, calcareous soils; 100-1000 m; Ont.; Ala., Ark., Conn., D.C., Ga., Ill., Ind., Iowa, Kans., Ky., Mich., Minn., Mo., N.Y., N.C., Ohio, Okla., Pa., Tenn., Vt., Va., W.Va., Wis.