Perennials, 60-150 cm (rhizomatous, sometimes stoloniferous as well). Stems (often reddish) erect, pilose to appressed-hairy proximally. Leaves mostly basal; opposite; petioles 2.5-10 cm; blades oblong-lanceolate or elliptic to ovate, 5-20 × 1.5-7 cm, bases cuneate, margins entire or serrulate, abaxial faces scabrous to hispidulous, gland-dotted (cauline usually smaller, rarely 2-4 pairs well developed). Heads 1-4(-12). Peduncles 1-14 cm. Involucres cylindric, 9-14 mm diam. Phyllaries 20-25, lanceolate, 4.5-7(-11) × 1.5-2.5 mm, (margins usually ciliate) apices acute to short-acuminate, abaxial faces glabrate to sparsely pilose (hairs moniliform), not gland-dotted. Paleae 5-7 mm, subentire to 3-toothed (shortly mucronate). Ray florets 8-14; laminae 18-22 mm (abaxial faces densely gland-dotted). Disc florets 50+; corollas 4.5-5.5 mm, lobes yellow; anthers dark brown or black, appendages yellow. Cypselae 3-4 (-5) mm, sparsely villous or glabrate; pappi of 2 aristate scales 1.5-2.8 mm plus 0-4 deltate scales 0.4-1 mm. 2n = 34 (subspecies unknown).