Perennials. Stems glabrous or glabrate. Leaves: petioles 3-10+ cm, seldom, if ever, winged; blades 4-20(-40+) × 2-12(25+) cm, larger usually deeply pinnately lobed, lobes 5-11. Cypselae ca. 3 mm, 4-6-angled or -ribbed. 2n = 30. Flowering Jul. Damp, shaded sites, calcareous soils; 10-300 m; Ala., Fla., Ga., Ky., Mo., Tenn.