Central Taxonomic Thesaurus

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  Animalia
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        Hexapoda
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                Neoptera
                    Coleoptera
                      Polyphaga
                        Scarabeiformia
                          Scarabaeoidea
                            Scarabaeidae
                              Aclopinae
                              Allidiostomatinae
                              Aphodiinae
                                Aegialiini
                                     Aegialia
                                     Caelius
                                     Micraegialia
                                Aphodiini
                                     Acrossus
                                     Agoliinus
                                     Aphodius
                                     Ballucus
                                     Blackburneus
                                     Calamosternus
                                     Caligodorus
                                     Cephalocyclus
                                     Chilothorax
                                     Cinacanthus
                                     Coelotrachelus
                                     Colobopterus
                                     Cryptoscatomaseter
                                     Dellacasiellus
                                     Dialytellus
                                     Dialytes
                                     Dialytodius
                                     Diapterna
                                     Drepanocanthoides
                                     Eupleurus
                                     Flaviellus
                                     Geomyphilus
                                     Haroldiellus
                                     Hornietus
                                     Irrasinus
                                     Labarrus
                                     Lechorodius
                                     Liothorax
                                     Luxolinus
                                     Maculaphodius
                                     Melinopterus
                                     Mendidius
                                     Merogyrus
                                     Nialaphodius
                                     Oscarinus
                                     Otophorus
                                     Oxyomus
                                     Paradalosus
                                     Pardalosus
                                     Planolinellus
                                     Planolinoides
                                     Planolinus
                                     Pseudagolius
                                     Rugaphodius
                                     Scabrostomus
                                     Schaefferellus
                                     Setodius
                                     Stenotothorax
                                     Strigodius
                                     Tetraclipeoides
                                     Teuchestes
                                     Xeropsamobeus
                                Didactyliini
                                     Aidophus
                                Eupariini
                                     Annegialia
                                     Aphotaenius
                                     Ataeniopsis
                                     Ataenius
                                     Euparia
                                     Euparixia
                                     Martineziana
                                     Pseudataenius
                                     Saprosites
                                Psammodiini
                                     Diastictus
                                     Geopsammodius
                                     Leiopsammodius
                                     Neopsammodius
                                     Odontopsammodius
                                     Parapsammodius
                                     Platytomus
                                     Pleurophorus
                                     Psammodius
                                     Rhyssemus
                                     Tesarius
                                     Trichiorhyssemus
                                     Aphonides
                              Cetoniinae
                                Cetoniini
                                  Cetoniina
                                     Chlorixanthe
                                     Heterocnemis
                                     Protaetia
                                     Dischista
                                     Oxythyrea
                                Cremastocheilini
                                  Cremastocheilina
                                     Cremastocheilus
                                     Genuchinus
                                     Lissomelas
                                     Psilocnemis
                                  Cremastochelinina
                                Gymnetini
                                     Amithao
                                  Gymnetina
                                     Cotinis
                                     Gymnetis
                                     Hologymnetis
                                Osmodermatini
                                     Osmoderma
                                     Rhomborrhina
                                Trichiini
                                     Gnorimella
                                     Trichiotinus
                                     Trigonopeltastes
                                Valgini
                                     Valgus
                                     Chlorixantha
                                     Cineretis
                              Dynamopodinae
                              Dynastinae
                                     Amblyodus
                                     Bothynus
                                Cyclocephalini
                                     Ancognatha
                                     Aspidolea
                                     Cyclocephala
                                     Dyscinetus
                                     Stenocrates
                                     Surutu
                                Dynastini
                                     Chalcosoma
                                     Dynastes
                                     Golofa
                                     Megasoma
                                Hexodontini
                                     Hemicyrthus
                                Oryctini
                                     Enema
                                     Strategus
                                     Xyloryctes
                                Pentodontini
                                     Anoplognatho
                                     Aphonus
                                     Coscinocephalus
                                     Euetheola
                                     Orizabus
                                     Oxygrylius
                                     Temnorhynchus
                                     Tomarus
                                Phileurini
                                     Archophileurus
                                     Hemiphileurus
                                     Phileurus
                                     Platyphileurus
                                     Phyllognathus
                                     Podalgus
                                     Spodistes
                                     Epicometis
                              Euchirinae
                                     Gymnopleurus
                                     Ligyrus
                                     Megathopa
                                     Melolontha
                              Melolonthinae
                                     Acoma
                                     Chaunocolus
                                     Chnaunanthus
                                Dichelonychini
                                     Coenonycha
                                     Dichelonyx
                                     Gymnopyge
                                Diplognathini
                                     Diplognatha
                                Diplotaxini
                                     Gymnoloma
                                Hopliini
                                     Eriesthis
                                     Hoplia
                                Lichniini
                                Macrodactylini
                                     Ceraspis
                                     Isonychus
                                     Macrodactylus
                                Melolonthini
                                     Amblonoxia
                                     Amphimallon
                                     Dinacoma
                                     Fossocarus
                                     Gronocarus
                                     Hypothyce
                                     Hypotrichia
                                     Phyllophaga
                                     Plectrodes
                                     Polyphylla
                                     Thyce
                                Oncerini
                                     Nefoncerus
                                     Oncerus
                                Pachydemini
                                     Phobetus
                                Podolasiini
                                     Podolasia
                                Sericini
                                     Maladera
                                     Nipponoserica
                                     Serica
                                     Metacatharsius
                              Orphninae
                              Pachypodinae
                                     Pachystethus
                                     Paracopris
                                     Parathyce
                              Phaenomeridinae
                                     Pinotus
                                     Plusiotis
                                     Polylamina
                                     Rhombonalia
                              Rutelinae
                                Adoretini
                                     Adoretus
                                Anomalini
                                     Anisoplia
                                  Anisopliina
                                     Anomalacra
                                  Anomalina
                                  Anomala
                                     Strigoderma
                                  Leptohopliina
                                     Leptohoplia
                                  Popilliina
                                     Popillia
                                     Heteronychus
                                     Macraspis
                                Rutelini
                                     Chrysina
                                     Chrysophora
                                     Cotalpa
                                     Parabyrsopolis
                                     Paracotalpa
                                     Parastasia
                                     Pelidnota
                                     Pseudocotalpa
                                     Rutela
                                     Spodochlamys
                                     Trizogeniates
                              Scarabaeinae
                                Ateuchini
                                     Ateuchus
                                Canthonini
                                     Anachalcos
                                     Boreocanthon
                                     Canthon
                                     Deltochilum
                                Coprini
                                     Canthidium
                                     Catharsius
                                     Copris
                                     Dichotomius
                                Oniticellini
                                     Euoniticellus
                                     Euonticellus
                                     Liatongus
                                     Oniticellus
                                     Tiniocellus
                                Onitini
                                     Onitis
                                Onthophagini
                                     Onthophagus
                                       Onthophagus (Digitonthophagus)
                                     Proagoderus
                                Phanaeini
                                     Coprophanaeus
                                     Phanaeus
                                Sisyphini
                                     Sisyphus